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Additional Information

Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Shi hongxiao conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, prepared figures and/or tables, authored or reviewed drafts of the paper, approved the final draft.

Hou xiangyang conceived and designed the experiments.

Wu xinghong conceived and designed the experiments, authored or reviewed drafts of the paper.

Song haiqun performed the experiments, contributed reagents/materials/analysis tools.

Yang tingting performed the experiments.

Data Deposition

The following information was supplied regarding data availability:

Data are not provided with this preprint.

Funding

This work was supported by the National Basic Research Program(grant number 2014CB138802) in China, the Project of Qinghai Science & Technology Department (2016-ZJ-Y01), the Open Project of State Key Laboratory of Plateau Ecology and Agriculture, Qinghai University (2016KF-06), Central Non-profit Research Institutes Fundamental Research Funds of China (1610332016003), Key Laboratory of Superior Forage Germplasm in the Qinghai-Tibetan Plateau (2017-ZJ-Y12). and the International Science and Technology Project of China (31560668). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.


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