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Additional Information

Competing Interests

There are no competing interests to declare.

Author Contributions

Hongdong Li conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper.

Qingsong Xu performed the experiments.

Yizeng Liang conceived and designed the experiments, contributed reagents/materials/analysis tools.


This work was financially supported by the National Nature Foundation Committee of P.R. China Grants No. 21075138. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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