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Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Wang Lu performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Hua Qingzhu performed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

Ma Yuewen performed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

Hu Guibing analyzed the data, reviewed drafts of the paper.

Qin Yonghua conceived and designed the experiments, analyzed the data, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Data Deposition

The following information was supplied regarding data availability:

The raw data has been supplied as a supplementary file.

Funding

This work was supported by the Science and Technology Planning Project of Guangdong Province (2010B020305007), Key Laboratory of Innovation and Utilization for Germplasm Resources in Horticultural Crops in Southern China of Guangdong Higher Education Institutes, the South China Agricultural University (No. KBL11008). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.


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