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Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Shambhu Paudel conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Juan C Levesque analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, reviewed drafts of the paper.

Camilo Saavedra analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Cristina Pita analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Prabhat Pal analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Human Ethics

The following information was supplied relating to ethical approvals (i.e., approving body and any reference numbers):

Department of National Parks and Wildllife Conservation (DNPWC) provided the approval to undertake this study in three river systems of Nepal. Reference number 353.

Funding

Ocean Park Conservation Foundation and Rufford Foundation provided funding. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.


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