Got Ph.D in Nagoya University, Japan, on 1995.
Postdoc in JT Biohistory Research Hall in Osaka, Japan (1996-1998) and Dalhousie University in Halifax, Canada (1998-2004).
Lecturer in Nagahama Institute for Bio-science and Technology in Nagahama (2004-2005).
Associate Professor in University of Tsukuba in Tsukuba (2006-2015).
Professor in University of Tsukuba in Tsukuba (2016-).