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Several papers and a website by Peters (see below) address several problems ignored by this manuscript/paper including:

  1. Basal pterosaur ichnites (they are known, have been published and non are quadrupedal, some are digitigade)
  2. The origin of pterosaurs (among bipedal fenestrasaurs each with uropatagia, trailing each hind limb. Objections to this scenario by Hone and Benton (2007, 2008)...
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Ichnology. Paleontology
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