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Indeed, the authors of the PLOS ONE paper discuss the unreliable nature of that Cretaceous "dung beetle" in the Discussion chapter (the fact that this fossil is doubtful was raised earlier).

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And yes, I would agree that dung beetles arose in Cretaceous -- their present distribution that is apparently associated with the break-up of Gondwana confirms it.

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Furthermore, the present distribution could indicate that they were affected by the break-up of Gondwana, but it could equally represent a case of pseudocongruence. A lot could of happened or affected their distribution in the last tens of millions of years since the break-up of Gondwana...

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