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Additional Information

Competing Interests

The author declares they have no competing interests.

Author Contributions

Christopher Hamm conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Field Study Permissions

The following information was supplied relating to field study approvals (i.e., approving body and any reference numbers):

This research was conducted under permit # 076-2010-SINAC as issued by the Minesterio de Ambiente, Energia y Telecomunicaciones (MINAE) of the Costa Rican government.

Data Deposition

The following information was supplied regarding the deposition of related data:

All data and code necessary to replicate the results presented in this paper will be deposited and made freely available on FigShare.


This project was made possible by a scholarship to the author from the Organization for Tropical Studies. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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