There is a typo in the equation at line 187; the Hill exponent n was inadvertently omitted. It should read:
E(D) = E0 + Emax ∙ Dn / ( Dn + 10nQ )
This and a few other minor errors will be corrected in the final peer-reviewed version, which has been accepted for publication in PeerJ (DOI 10.7717/peerj.195).
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