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Additional Information

Competing Interests

Yvan Richard and Edward R. Abraham are employees of Dragonfly Science and Lyndon Perriman is an employee of the New Zealand Department of Conservation.

Author Contributions

Yvan Richard analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Lyndon Perriman reviewed drafts of the paper, expertise; Data.

Chris Lalas reviewed drafts of the paper, expertise; Data.

Edward R. Abraham contributed reagents/materials/analysis tools, reviewed drafts of the paper.


This project was funded by the New Zealand Department of Conservation’s Conservation Services Programme ( project POP2011-09, partially through a levy on the quota holders of relevant commercial fish stocks. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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