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Additional Information

Competing Interests

The authors declare they have no competing interests.

Author Contributions

Sean D Smith performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Joseph K Kawash analyzed the data, reviewed drafts of the paper.

Andrey Grigoriev conceived and designed the experiments, analyzed the data, wrote the paper, reviewed drafts of the paper.


This work was in part supported by the NSF grant DBI-1126052 to AG. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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