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Additional Information

Competing Interests

The authors declare they have no competing interests.

Author Contributions

Chris J Barrett conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper, field work, creation of model, primary author.

Magnus L Johnson conceived and designed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper, help with model creation, comments which improved early drafts.

Susan L Hull conceived and designed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper, field work, help with draft and model creation.

Animal Ethics

The following information was supplied relating to ethical approvals (i.e., approving body and any reference numbers):

The Scarborough Campus (of Hull University) Ethics Committee approved the research.

An application was filled in and approved.

Funding

The authors declare there was no funding for this work.


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