A peer-reviewed article of this Preprint also exists.
This is an intersting work. Note that high polymorphism of rDNA has also been discovered in sing cells of ciliates (protist /single-cell eukaryote):
Gong J., Dong J., Liu X., Massana R. (2013) Extremely high copy numbers and polymorphisms of the rDNA operon estimated from single cell analysis of oligotrich and peritrich ciliates. Protist 164:369-379.
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