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Additional Information

Competing Interests

FC serves as an Academic Editor for PeerJ. He is the author of the book “The MPS Language Workbench, Volume I” and receives royalties from the sale of this book.

Author Contributions

Victoria M Benson performed the experiments, reviewed drafts of the paper.

Fabien Campagne conceived and designed the experiments, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.


This study was supported by the National Institutes of Health NIAID award 5R01AI107762-02 to FC. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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