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Additional Information

Competing Interests

Cynthia Parr is an employee of the National Museum of Natural History, Smithsonian Institution. Authors have no competing interests.

Author Contributions

Cynthia Parr conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper, reviewed drafts of the paper.

Craig R McClain conceived and designed the experiments, performed the experiments, reviewed drafts of the paper, logistics.


Support was provided by the Richard Lounsbery Foundation, Robert T. and Catherine T. MacArthur Foundation, and the National Evolutionary Synthesis Center (NESCent) NSF EF-0905606. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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