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Additional Information

Competing Interests

John R. Hutchinson is an Academic Editor for PeerJ. There are no other competing interests.

Author Contributions

Sophie Regnault performed the experiments, analyzed the data, wrote the paper, prepared figures and/or tables.

Andrew A Pitsillides analyzed the data, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

John R. Hutchinson conceived and designed the experiments, analyzed the data, reviewed drafts of the paper.

Animal Ethics

The following information was supplied relating to ethical approvals (i.e., approving body and any reference numbers):

Only cadaver material was used in this study; no ethical approval was required (the animals were not killed for this study).

Grant Disclosures

The following grant information was disclosed by the authors:

Leverhulme Trust grant number RPG-2013-108


This research was funded by a grant from the Leverhulme Trust (RPG-2013-108) to J.R.H. and A.P., and J.R.H.’s role was enabled by a Senior Research Fellowship awarded by the Royal Society Leverhulme Trust in 2012. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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