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Additional Information

Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Dong-Mei Wu performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables.

Jian-Xin Wang conceived and designed the experiments, reviewed drafts of the paper.

Xiao-Hui Liu performed the experiments.

Ying-Ping Fan reviewed drafts of the paper.

Ran Jiang performed the experiments.

Ming-Hua Liu performed the experiments.

Shuai-Bing Wang performed the experiments.

Xue-Zhu Liu reviewed drafts of the paper.

Data Deposition

The following information was supplied regarding data availability:

All the raw sequences in this study have been submitted to the NCBI-SRA public database ( under the ID: SRP104573 (all the twelve samples of the Changjiang Estuary).


This work was supported by the National Natural Science Foundation of China [grant numbers 31270160, J1310037] and the Natural Science Foundation of Zhejiang Province, China [grant number LY12C03003]. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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