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Additional Information

Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Long Wang conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, prepared figures and/or tables, authored or reviewed drafts of the paper, approved the final draft.

Sabin Saurav Pokharel analyzed the data, approved the final draft.

Fajun Chen conceived and designed the experiments, contributed reagents/materials/analysis tools, authored or reviewed drafts of the paper, approved the final draft.

Data Deposition

The following information was supplied regarding data availability:

The raw measurements are provided in the supplementary file 1 including associated raw values and four figures (figure 1 - figure 4).


This research was funded by the National Key Research and Development Program of China (2017YFD0200400), the Special Program for New Transgenic Variety Breeding of the Ministry of Science and Technology, China (2016ZX08012005), the State Public Industry (Agriculture) Research Project of China (201403031) and the Postgraduate Research &Practice Innovation Program of Jiangsu Province (KYCX17_0574). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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