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Additional Information

Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Xiaolin Chen conceived and designed the experiments, authored or reviewed drafts of the paper, approved the final draft.

Yuhao Sun performed the experiments.

Hong Liu contributed reagents/materials/analysis tools.

Song Liu analyzed the data.

Yukun Qin prepared figures and/or tables.

Pengcheng Li check the manuscript.

Data Deposition

The following information was supplied regarding data availability:

The research in this article did not generate any data or code (literature review).


This work was supported by the Commonweal Item of the State Oceanic Administration of the People’s Republic of China (201505033), NSFC-Shandong joint Fund (U1606403), Shandong Province Key Research and Development Project (2016YYSP010) and Qingdao People’s Livelihood Science and Technology Projects (16-6-2-41-nsh). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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