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Additional Information

Competing Interests

The authors declare no conflict of interest.

Author Contributions

Dongfeng Li conceived and designed the experiments, contributed reagents/materials/analysis tools, wrote the paper, reviewed drafts of the paper.

Songhua Wang conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper, prepared figures and/or tables.

Congshu Liao performed the experiments, prepared figures and/or tables.

Shaoyi Liu performed the experiments.

Fengling Li performed the experiments.

Wei Meng analyzed the data.

Animal Ethics

The following information was supplied relating to ethical approvals (i.e., approving body and any reference numbers):

All experiments were carried out in accordance with the university and national animal guidelines. The care and use of animals reported on in this study were approved by the Institutional Animal Care and Use Committee at South China Normal University and in accordance with National Institutes of Health Guidelines (scnu20070033).

Grant Disclosures

The following grant information was disclosed by the authors:

National Natural Science Foundation of China (31172092).


This work was supported by the National Natural Science Foundation of China (31172092). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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