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Additional Information

Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Emily Graham conceived and designed the experiments, analyzed the data, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Alex R. Crump conceived and designed the experiments, performed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

Charles T Resch performed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

Sarah Fansler performed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

Evan Arntzen conceived and designed the experiments, performed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

David Kennedy conceived and designed the experiments, performed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

Jim K Fredrickson conceived and designed the experiments, performed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

James C. Stegen conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

DNA Deposition

The following information was supplied regarding the deposition of DNA sequences:

Will be published upon acceptance in a peer-reviewed journal.

Data Deposition

The following information was supplied regarding data availability:

Will be published upon acceptance in a peer-reviewed journal.

Funding

This research was supported by the US Department of Energy (DOE), Office of Biological and Environmental Research (BER), as part of Subsurface Biogeochemical Research Program’s Scientific Focus Area (SFA) at the Pacific Northwest National Laboratory (PNNL). PNNL is operated for DOE by Battelle under contract DE-AC06-76RLO 1830. A portion of the research was performed using Institutional Computing at PNNL. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.


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