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Additional Information

Competing Interests

The authors declare they have no competing interests.

Author Contributions

Daniel J McGlinn conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper.

Michael W Palmer conceived and designed the experiments, performed the experiments, wrote the paper.

Data Deposition

The following information was supplied regarding the deposition of related data:


DJM received funding from the U.S. Environmental Protection Agency (EPA) under the Greater Research Opportunities Graduate Program. The U.S. EPA has not officially endorsed this publication, and the views expressed herein may not reflect the views of the Agency. MWP acknowledges support from NSF Grant Number EPS-0447262 and EPS-0919466, The Oklahoma State University College of Arts and Science, The Oklahoma Nature Conservancy, The Spatial and Environmental Information Clearinghouse, The Philecology Trust, The Oklahoma Water Resources Research Institute. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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