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Additional Information

Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Matthew C Metz conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Douglas W Smith conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Mark Hebblewhite conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Daniel R Stahler conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Animal Ethics

The following information was supplied relating to ethical approvals (i.e., approving body and any reference numbers):

University of Montana IACUC AUP 043-15

Field Study Permissions

The following information was supplied relating to field study approvals (i.e., approving body and any reference numbers):

National Park Service

Data Deposition

The following information was supplied regarding data availability:

The research in this article did not generate any raw data.

Funding

The authors received no funding for this work.


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