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Additional Information

Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Lucas R Forti conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper, prepared figures and/or tables.

William P Costa performed the experiments, contributed reagents/materials/analysis tools, reviewed drafts of the paper.

Lucas B Martins contributed reagents/materials/analysis tools, reviewed drafts of the paper.

Carlos H L Nunes-de-Almeida contributed reagents/materials/analysis tools, prepared figures and/or tables, reviewed drafts of the paper.

Luís Felipe Toledo conceived and designed the experiments, reviewed drafts of the paper.

Data Deposition

The following information was supplied regarding data availability:

Acoustic recordings used in the work are available in the Fonoteca Neotropical Jacques Vielliard, with collection numbers FNJV 10498, 10577, 12222–24, 12228, the Smithsonian Institution website (, and the personal collection of Ariovaldo A. Giaretta, which are detailed in the appendix of Martins & Giaretta (2012).


Funding was provided by Fundação de Amparo à Pesquisa do Estado de São Paulo (FAPESP) (fellowships (2013/09964-2, 2013/21519-4 and 2013/02219-0)); and Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) (fellowships (302589/2013-9, 152548/2011-4) and a research grant (405285/2013-2) to LFT). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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