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Additional Information

Competing Interests

There are no competing interests.

Author Contributions

Grant Harris conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper.

Sean Farley conceived and designed the experiments, contributed reagents/materials/analysis tools, wrote the paper.

Gareth J Russell conceived and designed the experiments, analyzed the data, wrote the paper.

Matthew J Butler conceived and designed the experiments, analyzed the data, wrote the paper.

Jeff Selinger conceived and designed the experiments, contributed reagents/materials/analysis tools, wrote the paper.

Animal Ethics

The following information was supplied relating to ethical approvals (i.e., approving body and any reference numbers):

All field and capture methods were approved by Alaska Department of Fish and Game, Animal Care and Use Committee, Assurance No. 06-03.

Funding

The Alaska Department of Fish and Game, the USDA Forest Service, National Park Service and U.S. Fish and Wildlife Service provided logistical and financial backing. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.


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