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Additional Information

Competing Interests

Dorothy V Bishop is an Academic Advisor and an Academic Editor for PeerJ.

Author Contributions

Dorothy V Bishop conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper, programming simulation.

Paul A Thompson performed the experiments, analyzed the data, prepared figures and/or tables, reviewed drafts of the paper, programming simulation.

Data Deposition

The following information was supplied regarding the deposition of related data:

Code for Ghostphack is provided in Appendix 2. We are still developing this code and plan a further paper based on this. We will deposit a final version on a public database such as Figshare or Open Science Framework.

Funding

The first author is supported by a Wellcome Trust Principal Research Fellowship. Both authors are funded by Wellcome Trust Programme Grant number 082498/Z/07/Z. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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