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Additional Information

Competing Interests

Georgia C.A. French, Joanna K. Bluemelare, and David Rowat are employees of Marine Conservation Society Seychelles.

Author Contributions

Georgia C.A. French conceived and designed the experiments, performed the experiments, analyzed the data, wrote the paper.

Joanna K Bluemel editorial input, statistical advice.

David Rowat contributed reagents/materials/analysis tools, editorial input.

Animal Ethics

The following information was supplied relating to ethical approvals (i.e., approving body and any reference numbers):

No approval required for the type of research undertaken.


Funding for this study was provided by the Marine Conservation Society Seychelles, the Save Our Seas Foundation and the Global Environment Facility (Administered by the United Nations Development Programme). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.

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