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Born in Hamilton, Ontario, Canada. Grew up in Patras, Greece. Biology degree from the University of Patras. Ph.D. with Herbert Jäckle at the Max-Planck Insitute (Göttingen) and John P. Phillips at the University of Guelph on the genetic regulation of reactive oxygen species in Drosophila. Post-doc with Tracey Rouault at the National Institute of Child Health and Human Development, in Bethesda, Maryland on iron metabolism. First post at Queen Mary, University of London. Metals in Biology.