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Stebner et al. (2015) recently described a new subfamily of fossil Psychodidae. However, the establishment of a new subfamily Protopsychodinae Stebner et al., 2015 with the type genus being Datzia is invalid under ICZN rules, article 29.1, because the subfamily name must be formed using the stem of the type genus name. Furthermore, Protopsychoda Azar et al., 1999 is a valid genus described within the subfamily Psychodinae which would make Protopsychodinae (if validly established with Protopsychoda as the type genus) a junior synonym of Psychodinae (ICZN, article 23). The name Protopsychodinae is unavailable, and instead the new subfamily name Datziinae is established, based on Datzia Stebner et al., 2015 as the type genus. Accordingly, the new subfamily name is herein proposed to include the type genus Datzia and the genus Mandalayia Stebner et al., 2015.

Material and Methods

The electronic version of this article in Portable Document Format (PDF) will represent a published work according to the International Commission on Zoological Nomenclature (ICZN), and hence the new names contained in the electronic version are effectively published under that Code from the electronic edition alone. This published work and the nomenclatural acts it contains have been registered in ZooBank, the online registration system for the ICZN. The ZooBank LSIDs (Life Science Identifiers) can be resolved and the associated information viewed through any standard web browser by appending the LSID to the prefix “”. The LSID for this publication is: The online version of this work is archived and available from the following digital repositories: PeerJ, PubMed Central and CLOCKSS.

Systematic Palaeontology

Order: Diptera Linnaeus, 1758
Family: Psychodidae Newman, 1834

Datziinae new subfamily (replacing Protopsychodinae Stebner et al., 2015, unavailable under the ICZN Code)

Type-genus: Datzia Stebner et al., 2015

Type-species: Datzia bispina

Material: Holotype male SMF Be 2379 and 7 male Paratypes; SMF Be 2376 Paratypes 6 males. Deposited at the Senckenberg Forschungsinstitut und Naturmuseum, Frankfurt, Germany.

Type-locality: Hukawng Valley of the northern state of Kachin.

Stratigraphic horizon: 98.79 ± 0.62 million years, Upper Cretaceous (earliest Cenomanian) (Shi et al., 2012).

Diagnosis: unchanged (see Stebner et al., 2015, p. 6)


The authors would like to thank Christian Kammerer (Museum für Naturkunde, Berlin) and Matthew E. Clapham (University of California, Santa Cruz) for pointing out the nomenclatural inconsistency in our previous paper, Richard Pyle (ZooBank) for reviewing the manuscript and providing very helpful advice for this work, and Jacqueline Thai (Head of Publishing Operations, PeerJ) for her constant efforts.

Additional Information and Declarations

Competing Interests

The authors declare there are no competing interests. Mónica M Solórzano Kraemer is an employee of the Senckenberg Forschungsinstitut und Naturmuseum, Sektion Mikropaläontologie I-Bernstein.

Author Contributions

Frauke Stebner, Mónica M. Solórzano Kraemer, Sergio Ibáñez-Bernal and Rüdiger Wagner conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, reviewed drafts of the paper.

New Species Registration

The following information was supplied regarding the registration of a newly described species:

1. This published work and the nomenclatural acts it contains have been registered in ZooBank.

The LSID for this publication is:

2. New subfamily Datziinae:


This research was possible with funding No. RU665/10-1 from the German Research Foundation (DFG), and a postdoctoral fellowship to Mónica M. Solórzano Kraemer, No. SO894/3-1 from the German Research Foundation (DFG). Sergio Ibáñez-Bernal was supported by the Project INECOL-10816. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.