Buchanan Professor of Genetics at the University of Edinburgh & a Fellow of the Royal Societies of London & Edinburgh. Director of the Wellcome Trust Centre for Cell Biology from 1999 to 2011. Governor of the Wellcome Trust from 2000-10 & Deputy Chairman from 2007. Currently trustee of Cancer Research UK & the Kirkhouse Trust. Awards include Howard Hughes International Fellowship, the Louis-Jeantet Prize for Medicine, the Gairdner Foundation prize and the Gabor & GSK Medals of the Royal Society.